GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	September 11, 2024

Displaying entries **3526 - 3550** of **5867** in total

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GlyTouCan ID ▲	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G59149SL	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Sialylated LDN	Linkage defined saccharide	2159.8	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59155GF	Fuc(a1-2)Gal(b1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	H antigen (type 3) O-Glycan core 2 Polysialic acid	Linkage defined saccharide	1185.43	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59168YE	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(?1-	GM4	Linkage defined saccharide	674.24	Amniota Bifurcata Bilateria Boreoeutheria Calloselasma rhodostoma Calloselasma Catarrhini Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lepidosauria Mammalia Metazoa Opisthokonta Primates Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G59185WM	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1437.51	Amniota Bacteria Bilateria Boreoeutheria Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Mammalia Metazoa Opisthokonta Primates Pseudomonadota Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59229NY	HexNAc(b1-3/4)Gal(b1-3)[GalNAc(b1-4)GlcNAc(b1-6)]GalNAc(a1-	LDN LacdiNAc O-Glycan core 2	Linkage defined saccharide	992.38	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59234CC	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)[GlcNAc(b1-2)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	1915.69	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59277TZ	GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1868.7	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59297UK	Neu5Ac(?2-?)Gal(?1-?)[Fuc(?1-?)]GlcNAc(?1-?)Man(a1-3/6)[Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2514.9	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59303WP	Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)[Gal(?1-?)GlcNAc(?1-?)]Man(a1-3)[Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2443.9	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59306VK	GlcNAc(b1-3)[Neu5Ac(a2-6)]Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1373.51	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59311AO	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan high mannose	Linkage defined saccharide	1396.49	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acremonium Amniota Amphiesmenoptera Amphipyrinae Anthemideae Artemisia vulgaris Artemisia Artemisiinae Arthropoda Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asteroideae BOP clade Batoidea Bilateria Blastocrithidia Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chondrichthyes Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Discoba Ditrysia Ecdysozoa Elasmobranchii Embryophyta Endopterygota Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fungi Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Herpetomonas samuelpessoai Herpetomonas Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hypocreales incertae sedis Hypocreales Hypocreomycetidae Insecta Kinetoplastea Laurasiatheria Leishmania adleri Leishmania mexicana species complex Leishmania mexicana Leishmania Leishmania Leishmaniinae Lepidoptera Leptomonas samueli Leptomonas Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Petrosaviidae Pezizomycotina Phaseoleae Phaseolus vulgaris Phaseolus Poaceae Poales Primates Protostomia Pterygota Ricinus communis Ricinus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Schizotrypanum Simiiformes Sordariomycetes Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Strigomonadinae Strigomonas culicis Strigomonas Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Tracheophyta Trypanosoma conorhini Trypanosoma cruzi Trypanosoma dionisii Trypanosoma with unspecified subgenus Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata Viridiplantae asterids campanulids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade leotiomyceta lizard Leishmania rosids saccharomyceta sordariomyceta
G59334JE			Base composition with linkage	2290.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G59354CW			Monosaccharide composition with linkage	2181.76	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59358BQ	Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(?1-?)]GlcNAc(b1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1421.52	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59361OW	GalNAc(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	2071.78	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59411JK		N-Glycan core basic	Linkage defined saccharide	2264.83	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59419HU	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)][Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc	CA19-9 antigen Sialyl Lewis a Slea	Linkage defined saccharide	1435.5	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59456VR	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2192.81	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59471JC	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2882.05	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59471TH	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1519.57	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Coturnix japonica Coturnix Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Perdicinae Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G59508TM	Gal(b1-3)GalNAc(b1-4)Gal3S(b1-4)Glc(b1-	Glucosylceramide	Linkage defined saccharide	787.21	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59513YA	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Acalypheae Acalyphoideae Acalyptratae Amniota Amphiesmenoptera Arthropoda Articulavirales Artiodactyla Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Insthoviricetes Kinetoplastea Laurasiatheria Lepidoptera Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Orthomyxoviridae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sarcopterygii Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids unclassified Orthomyxoviridae unidentified influenza virus
G59522PY	GlcNAc(?1-?)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1275.46	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59536GA			Glycosidic topology	2296.82	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G59575FI		N-Glycan hybrid N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1971.65	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms