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MIRAGE
G64973KT

Summary
GlyTouCan ID
G64973KT
IUPAC Condensed
Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-
Motifs
N-acetyllactosamine O-Glycan core 2 Type 2 LN
Subsumption Level
Fully-defined saccharide
Monoisotopic Mass
748.27
GlyConnect structure
GlyGen
ChEBI
PubChem Compound
PubChem Substance
CarbBank
3D Structures
GLYCAM
DGalpb1-3[DGalpb1-4DGlcpNAcb1-6]DGalpNAca1-OH
GlycoShape
G64973KT
Displaying 1 entry
Archetype α β
G96017QA
G64973KT
G01759NM
Organisms
Displaying entries 1 - 5 of 7 in total
Organisms Evidence
synthetic construct
Ovis aries (sheep)
Mus musculus (house mouse)
Bos taurus (domestic cattle)
Rattus norvegicus (Norway rat)
Taxonomic Hierarchy
GlycoGene Database (GGDB)
Displaying entries 1 - 5 of 8 in total
Gene Symbol Donor Acceptor Reducing terminal(Acceptor) Product Reducing terminal(Product) Reference
ST3GAL4 CMP-Neu5Ac
G64973KT
O-glycan Synthesis
G11457RF
O-glycan Synthesis
FUT6 GDP-Fuc
G64973KT
O-glycan Synthesis
G17752YB
O-glycan Synthesis
B4GALT4 UDP-Gal
G00033MO
pNP
G64973KT
pNP
B4GALT1 UDP-Gal
G00033MO
O-glycan Synthesis
G64973KT
O-glycan Synthesis
A4GNT UDP-GlcNAc
G64973KT
[alpha]pNP
G32910OR
[alpha]pNP
Displaying 1 entry
Gene Symbol Donor Acceptor Reducing terminal(Acceptor) Reference
B4GALT1 UDP-Gal
G64973KT
O-glycan Synthesis
KEGG BRITE Database
KEGG ID
Enzyme
References
Reactions
Displaying 1 entry
KEGG Reaction Enzyme Acceptor Product
R07617
Core Protein
Displaying entries 11 - 20 of 25 in total
UniProt ID Protein Name Reference Source
P13473 Lysosome-associated membrane glycoprotein 2
P14780 Matrix metalloproteinase-9
P15941 Mucin-1
P20239 Zona pellucida sperm-binding protein 2
P21754 Zona pellucida sperm-binding protein 3
P35253 Envelope glycoprotein
P80195 Glycosylation-dependent cell adhesion molecule 1
P98088 Mucin-5AC
Q02817 Mucin-2
Q14242 P-selectin glycoprotein ligand 1
GlycomeAtlas

Tissues in which this glycan is expressed are highlighted in yellow.

Danio rerio
Brain Gill Heart Intestine Intestine Liver Ovary Skin Testis
Literature
Tissue
  • heart
  • intestine
  • testis
Reference
Yamakawa N et al. Systems glycomics of adult zebrafish identifies organ-specific sialylation and glycosylation patterns. Nat Commun. 2018;9(1):4647. Published 2018 Nov 7. doi:10.1038/s41467-018-06950-3
DOI
10.1038/s41467-018-06950-3
Localization sites organized by hierarchy by Uber-anatomy ontology (UBERON)
Sequence Descriptors
GlycoCT
RES
1b:a-dgal-HEX-1:5
2s:n-acetyl
3b:b-dgal-HEX-1:5
4b:b-dglc-HEX-1:5
5s:n-acetyl
6b:b-dgal-HEX-1:5
LIN
1:1d(2+1)2n
2:1o(3+1)3d
3:1o(6+1)4d
4:4d(2+1)5n
5:4o(4+1)6d
WURCS
WURCS=2.0/3,4,3/[a2112h-1a_1-5_2*NCC/3=O][a2112h-1b_1-5][a2122h-1b_1-5_2*NCC/3=O]/1-2-3-2/a3-b1_a6-c1_c4-d1
Literature
Displaying entries 11 - 20 of 46 in total
PubMed ID Title First Author Publication Date Source
11323440 Molecular cloning and expression of a novel human beta-Gal-3-O-sulfotransferase that acts preferentially on N-acetyllactosamine in N- and O-glycans Suzuki A 2001 Jun 29
  • PubChem
11123894 O-Glycan Analysis of Natural Human Neutrophil Gelatinase B Using a Combination of Normal Phase- HPLC and Online Tandem Mass Spectrometry: Implications for the Domain Organization of the Enzyme Mattu TS 2000 Dec 01
  • GlyGen
  • PubChem
10430883 Expression cloning of a human alpha1, 4-N-acetylglucosaminyltransferase that forms GlcNAcalpha1-->4Galbeta-->R, a glycan specifically expressed in the gastric gland mucous cell-type mucin Nakayama J 1999 Aug 03
  • PubChem
9492296 Structural study of the O-linked sugar chains of human leukocyte tyrosine phosphatase CD45 Furukawa K 1998 Jan 15
  • GlyGen
  • PubChem
9530955 Purification and characterization of the MUC1 mucin-type glycoprotein, epitectin, from human urine: structures of the major oligosaccharide alditols Bhavanandan VP 1998 Jan
  • GlyGen
  • PubChem
9857011 Synthesis of poly-N-acetyllactosamine in core 2 branched O-glycans. The requirement of novel beta-1,4-galactosyltransferase IV and beta-1,3-n-acetylglucosaminyltransferase Ujita M 1998 Dec 25
  • PubChem
9180155 Isolation and characterization of ovarian cancer antigen CA 125 using a new monoclonal antibody (VK-8): identification as a mucin-type molecule Lloyd KO 1997 May 29
  • GlyGen
  • PubChem
9449027 Carbohydrate and peptide structure of the alpha- and beta-subunits of human chorionic gonadotropin from normal and aberrant pregnancy and choriocarcinoma Elliott MM 1997 Aug
  • GlyGen
  • PubChem
8702529 Structures of the O-Glycans on P-selectin Glycoprotein Ligand-1 from HL-60 Cells Wilkins PP 1996 Aug 02
  • GlyGen
  • PubChem
8323299 Assignment of O-Glycan Attachment Sites to the Hinge-like Regions of Human Lysosomal Membrane Glycoproteins Lamp-1 and Lamp-2 Carlsson SR 1993 Jul
  • GlyGen
  • PubChem
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Acknowledgements

Supported by JST NBDC Grant Number JPMJND2204

Partly supported by NIH Common Fund Grant #1U01GM125267-01

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Last updated: August 4, 2025