GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	September 11, 2024

Displaying entries **276 - 300** of **5867** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▼	Monoisotopic Mass	Organisms
G64973KT	Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN	Linkage defined saccharide	748.27	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovis aries Ovis Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G65562ZE	Neu5Ac(a2-3)Gal(b1-3)GalNAc(a1-	Sialyl T antigen	Linkage defined saccharide	674.24	Actinopteri Actinopterygii Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G49299ZV	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan hybrid N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2094.76	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G77824XV	Neu5Ac(a2-3)Gal(?1-		Linkage defined saccharide	471.16	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G74621DY	Gal(b1-4)Glc(?1-		Linkage defined saccharide	342.12	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00040MO	GalNAc(a1-3)GalNAc	O-Glycan core 5 fuzzy	Linkage defined saccharide	424.17	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G00042MO	GlcNAc(b1-6)GalNAc	O-Glycan core 6 Fuzzy	Linkage defined saccharide	424.17	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00044MO	GalNAc(a1-6)GalNAc	O-Glycan core 7 fuzzy	Linkage defined saccharide	424.17	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00070MO	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	707.25	Amniota Artiodactyla Bacteria Betaproteobacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neisseria gonorrhoeae FA19 Neisseria gonorrhoeae Neisseria meningitidis Neisseria Neisseriaceae Neisseriales Opisthokonta Pecora Primates Pseudomonadota Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G90889KG	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Actinopteri Actinopterygii Alphavirus Alsuviricetes Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops aethiops Chlorocebus aethiops Chlorocebus Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Laurasiatheria Mammalia Martellivirales Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Orthornavirae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rattus Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Sindbis virus Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms unclassified entries unclassified sequences unidentified
G00140YZ	Gal(b1-4)GlcNAc(b1-3/6)[Fuc(a1-2)Gal(b1-3/6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	896.35	Amniota Bilateria Boreoeutheria Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Macaca radiata Macaca Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00212ZT	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	1355.46	Adenoviridae Amniota Artiodactyla Bamfordvirae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Magnoliopsida Mammalia Mastadenovirus Mesangiospermae Metazoa Muroidea Myomorpha Opisthokonta Pecora Pentapetalae Preplasmiviricota Primates Rodentia Rowavirales Ruminantia Sarcopterygii Simiiformes Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Streptophyta Streptophytina Tectiliviricetes Teleostomi Tetrapoda Theria Tracheophyta Varidnaviria Vertebrata Viridiplantae Viruses asterids cellular organisms eudicotyledons lamiids unclassified Mastadenovirus unidentified adenovirus
G00470UO	GalNAc(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	2071.78	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00633GI	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-3/6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)]Man(b1-4)GlcNAc	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1802.64	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00675GF	Fuc(a1-2)Gal(b1-3/4)[Fuc(a1-3/4)]GlcNAc(b1-3/6)[Fuc(a1-2)Gal(b1-3/4)GlcNAc(b1-3/6)]Gal(b1-3/4)GlcNAc(b1-3)Gal(b1-3)[Gal(b1-6)]GalNAc(?1-	Blood group H (type 1) Blood group H (type 2) Blood group H (type 2) I' antigen Leb Leb Lewis b Lewis b Ley O-Glycan core 1 fuzzy Type 1 LN2	Linkage defined saccharide	2078.77	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00711QT			Linkage defined saccharide	475.06	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00688TE			Linkage defined saccharide	433.05	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00868ZN	Gal(b1-?)GlcNAc(b1-2)Man(a1-3)[Gal(b1-?)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	1640.59	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00883ZT	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan high mannose	Linkage defined saccharide	1558.54	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acremonium Amniota Amphiesmenoptera Amphipyrinae Arthropoda Artiodactyla Ascomycota BOP clade Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chordata Craniata Crithidia fasciculata Crithidia Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Discoba Ditrysia Ecdysozoa Embryophyta Endopterygota Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Felidae Feliformia Felinae Felis catus Felis Fungi Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hypocreales incertae sedis Hypocreales Hypocreomycetidae Insecta Kinetoplastea Laurasiatheria Leishmaniinae Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Petrosaviidae Pezizomycotina Poaceae Poales Primates Protostomia Pterygota Ricinus communis Ricinus Rodentia Ruminantia Sarcopterygii Simiiformes Sordariomycetes Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trypanosomatida Trypanosomatidae Vertebrata Viridiplantae cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade leotiomyceta rosids saccharomyceta sordariomyceta
G00978TF	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1275.46	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01028CW	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-3/6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2153.8	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muroidea Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01040ZV	Gal(b1-3)[Neu5Ac(a2-6)]GalNAc	O-Glycan core 1 fuzzy	Linkage defined saccharide	674.24	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01031EZ	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)[Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2079.76	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01347XI	Gal(a1-4)Gal(b1-4)GlcNAc		Linkage defined saccharide	545.2	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01329EH	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc	Lex-Lex	Linkage defined saccharide	1364.5	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms